Lexikon

Humus is the biochemical substance that makes the upper layers of the soil become dark. It is colored dark brown to black. Humus is difficult to see in isolation because it binds with larger mineral and organic particles. Humus provides soil with a number of very important benefits:

• It enhances a soil's ability to hold and store moisture.
• It reduces the eluviation of soluble nutrients from the soil profile.
• It is the primary source of nutrients required by plants for their nutrition.
• It improves soil structure which is necessary for plant growth.

Humus refers to organic matter that has decomposed to a point where it is resistant to further breakdown or alteration. Humic acids and fulvic acids are important constituents of humus and typically form from plant residues like foliage, stems and roots. After death, the plant residues begin to decay, starting the formation of humus. Humus formation involves changes within the soil and plant residue, there is a reduction of water soluble constituents including cellulose and hemicellulose; as the residues are deposited and break down, humin, lignin and lignin complexes accumulate within the soil; as microorganisms live and feed on the decaying plant matter, an increase in proteins occurs.

Lignin is resistant to breakdown and accumulates within the soil; it also chemically reacts with amino acids which add to its resistance to decomposition, including enzymatic decomposition by microbes. Fats and waxes from plant matter have some resistance to decomposition and persist in soils for a while. Clay soils often have higher organic contents that persist longer than soils without clay. Proteins normally decompose readily, but when bound to clay particles they become more resistant to decomposition. Clay particles also absorb enzymes that would break down proteins. The addition of organic matter to clay soils, can render the organic matter and any added nutrients inaccessible to plants and microbes for many years, since they can bind strongly to the clay. High soil tannin polyphenol content from plants can cause nitrogen to be sequestered by proteins or cause nitrogen immobilization, also making nitrogen unavailable to plants.

Humus formation is a process dependent on the amount of plant material added each year and the type of base soil; both are affected by climate and the type of organisms present. Soils with humus can vary in nitrogen content but have 3 to 6 percent nitrogen typically; humus, as a reserve of nitrogen and phosphorus, is a vital component affecting soil fertility.Humus also absorbs water, acting as a moisture reserve, that plants can utilize; it also expands and shrinks between dry and wet states, providing pore spaces. Humus is less stable than other soil constituents, because it is affected by microbial decomposition, and over time its concentration decreases without the addition of new organic matter. However, some forms of humus are highly stable and may persist over centuries if not millennia: they are issued from the slow oxidation of charcoal, also called black carbon, like in Amazonian Terra preta or Black Earths produced by ancient Indians.